Wheeler Shale
The Wheeler Shale (named by Charles Walcott) is a Cambrian (c. 507 Ma) fossil locality world famous[1] for prolific agnostid and Elrathia kingii trilobite remains (even though many areas are barren of fossils)[2] and represents a Konzentrat-Lagerstätten. Varied soft bodied organisms are locally preserved, a fauna (including Naraoia, Wiwaxia and Hallucigenia) and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale.[3] As such, the Wheeler Shale also represents a Konservat-Lagerstätten.[4]
Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms 490 to 610 m (1,600 to 2,000 ft) of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America.[5]
At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thin, flaggy limestone.[6] The Wheeler Formation (although the Marjum & Weeks Formations are missing) extends into the Drum Range, northwest of the House Range where similar fossils and preservation are found.[6]
Taphonomy & Sedimentology
Detailed work recognises a number of ~10 m thick lagerstätten sequences in the formation, each of which formed at a sea-level high stand[7] in deep water.[8] The lagerstätte were deposited by turbitides and mudslides onto an oxygenated sea floor.[7]
Stratigraphy
The Wheeler Shale spans the Ptychagnostus atavus[9] and uppermost-Middle Cambrian Bolaspidella trilobite zones (See House Range) for full stratigraphy).
Fauna
Incomplete list of the fauna of the Wheeler Shale:[6][10][11][12][13][14][15] (Note: the preservation of hard bodied trilobite remains and soft bodied animals seems to be mutually exclusive within particular horizons.)[4][16]
- Naraoia compacta - naraoiid nectaspid
- Hypagnostus parvifrons - agnostid
- Peronopsis amplaxis - peronopsid agnostid
- Peronopsis bidens
- Peronopsis fallax
- Peronopsis gaspensis
- Peronopsis intermedius
- Peronopsis interstrictus
- Peronopsis montis
- Peronopsis segmentis
- Ptychagnostus atavus (= Acidusus atavus) - ptychagnostid agnostid
- Ptychagnostus germanus
- Ptychagnostus gibbus
- Ptychagnostus intermedius
- Ptychagnostus michaeli
- Ptychagnostus occultatus
- Ptychagnostus seminula
- Glyphaspis concavus - asaphid
- Bathyuriscus fimbriatus - dolichometopid corynexochid
- Bathyuriscus sp.
- Kootenia sp. - dorypygid corynexochid, perhaps a synonym of Olenoides
- Olenoides expansus - dorypygid corynexochid
- Olenoides nevadensis
- Olenoides serratus
- Tonkinella breviceps
- Zacanthoides divergens - zacanthoidid corynexochid
- Zacanthoides sp.
- Altiocculus harrisi - ptychopariid (specific name may be confused with Alokistocare)
- Alokistocare harrisi - alokistocarid ptychopariid
- Asaphiscus wheeleri - ptychopariid; second-most common species in the formation
- Bathyocos housensis - ptychopariid
- Bolaspidella drumensis
- Bolaspidella housensis
- Bolaspidella sp.
- Bolaspidella wellsvillensis
- Brachyaspidion microps
- Brachyaspidion sulcatum
- Cedaria minor - known from the Warrior Formation
- Elrathia kingii - alokistocarid ptychopariid
- Elrathia sp.
- Elrathina wheeleri = Ptychoparella wheeleri? - ptychopariid
- Jenkinsonia varga
- Modocia brevispina
- Modocia laevinucha
- Modocia typicalis
- Ptychoparella sp. - ptychopariid
- Ptychoparella wheeleri
- Spencella sp. - ptychopariid
- Castericystis sprinklei - carpoid
- Castericystis sp.
- Cothurnocystis sp. - stylophoran
- Ctenocystis sp. - ctenocystoid
- Gogia spiralis - eocrinoid
- Eocrinoid holdfasts believed to belong to Gogia spiralis; may belong to other species
Unclassified
Gallery
References
- ^ Johnson, Kirk; Troll, Ray (2007), Cruising the fossil freeway: An epoch tale of a scientist and an artist on the ultimate 5,000-Mile paleo road trip, Golden, CO.: Fulcrum Publishing, ISBN 978-1-55591-451-6
- ^ Robert R. Gaines; Mary L. Droser (2003), "Paleoecology of the familiar trilobite Elrathia kingii: An early exaerobic zone inhabitant" (pdf), Geology 31: 941–4, doi:10.1130/G19926.1, http://earthsciences.ucr.edu/docs/Gaines&Droser_2003.pdf
- ^ Robert R. Gaines; Derek E.G. Briggs ; Zhao Yuanlong (2008), "Cambrian Burgess Shale–type deposits share a common mode of fossilization", Geology 36: 755–758, doi:10.1130/G24961A.1
- ^ a b Gaines, R; Kennedy, M; Droser, M (2005), "A New Hypothesis for Organic Preservation of Burgess Shale Taxa in the Middle Cambrian Wheeler Formation, House Range, Utah", Palaeogeography Palaeoclimatology Palaeoecology 220: 193–205, doi:10.1016/j.palaeo.2004.07.034
- ^ Robison, R.A. (1964), "Late Middle Cambrian faunas from western Utah", Journal of Paleontology 38 (3): 510–566, http://jpaleontol.geoscienceworld.org/cgi/content/abstract/38/3/510
- ^ a b c Gunther, L.F.; Gunther, V.G. (1981), "Some Middle Cambrian Fossils of Utah", Brigham Young University Geology Studies 28: 1–81
- ^ a b Brett, C. E.; Allison, P. A.; Desantis, M. K.; Liddell, W. D.; Kramer, A. (2009). "Sequence stratigraphy, cyclic facies, and lagerstätten in the Middle Cambrian Wheeler and Marjum Formations, Great Basin, Utah". Palaeogeography Palaeoclimatology Palaeoecology 277: 9–33. doi:10.1016/j.palaeo.2009.02.010. edit
- ^ Halgedahl, S. L.; Jarrard, R. D.; Brett, C. E.; Allison, P. A. (2009). "Geophysical and geological signatures of relative sea level change in the upper Wheeler Formation, Drum Mountains, West-Central Utah: A perspective into exceptional preservation of fossils". Palaeogeography Palaeoclimatology Palaeoecology 277 (1–2): 34–56. doi:10.1016/j.palaeo.2009.02.011. edit
- ^ Lieberman, B. S. (2003). "A New Soft-Bodied Fauna: the Pioche Formation of Nevada". Journal of Paleontology 77 (4): 674–690. doi:10.1666/0022-3360(2003)077<0674:ANSFTP>2.0.CO;2. ISSN 0022-3360. edit
- ^ Briggs, D.E.G.; Robison, R.A. (1984), Exceptionally preserved nontrilobite arthropods and Anomalocaris from the Middle Cambrian of Utah, The Paleontological Institute, The University of Kansas, http://kuscholarworks.ku.edu/dspace/handle/1808/3656
- ^ Photos of Wheeler Shale fossils from UC Berkeley
- ^ Rigby, J.K. (1978), "Porifera of the Middle Cambrian Wheeler Shale, from the Wheeler Amphitheater, House Range, in western Utah.", Journal of Paleontology 52 (6): 1325–1345
- ^ Utah's Cambrian Life from University of Kansas Natural History Museum
- ^ Cambrian fossils from Utah by the University of Utah
- ^ Comprehensive treatment from The Virtual Fossil Museum
- ^ Gaines, Robert R.; Droser, Mary L.; Kennedy, Martin J. (2001), "Taphonomy of soft-bodied preservation and ptychopariid Lagerstätte in the Wheeler Shale (Middle Cambrian), House Range, USA; controls and implications.", PaleoBios 21 (Suppl.2): 1–55, http://www.ucmp.berkeley.edu/napc/abs9.html
Modes of preservation in the Cambrian
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Exceptional |
Phosphate
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Carbonaceous film
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Casts and moulds
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Conventional |
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